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You can enable tree security on an object by setting a value for the security_tree_object and tree_assignment_object_name attributes with the Vault API’s Execute MDL Script endpoint. You can also execute MDL commands with Vault Toolbox.

For example, the following code enables tree security on the campaign__c custom object:

ALTER Object campaign__c ( security_tree_object('Object.security_tree__c'), tree_assignment_object_name('user_tree_assignment') );